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We need to find something that can breach the field. Something marked by death.
―A captive Talwyn Apogee to Ratchet  [QfB]
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Pirate skulls were the removed heads of undead pirates. After killing them, Ratchet could pick up pirate skulls with his OmniWrench Millennium 12 and throw them as long-range weapons.

They were used to destroy undead force fields that could only be deactivated by something marked by death, and dealt damage to Captain Darkwater when the boss was in his giant ghost form.

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